Scientific name: Hayata glandulifera Aver.
English name:
Vietnamese name:
Other name:

Hayata glandulifera Aver.

Plantae terrestres vel lithophilae, ad 15 cm alt., foliis (2)3(4), 4–8 cm lg., 1.5–3 cm lt. Scapus ad 5 cm lg., dense tomentosus, floribus albis 1–3, 9–10 mm lg., 6–7 mm lt. Sepala 8–9 mm lg., 3.5-4 mm lt. Petala 9–10 mm lg., 4–4.5 mm lt. Hypochilium 4-4.5 mm lg., 3–4 mm lt., glandulis capitatis magnis 6–7 ab utroque latere congestis. Mesochilium 1.5-2 mm lg. Epichilium bilobum, lobis 4.5–5.5 mm lg., 4–5 mm lt., irregulariter dentatis. Columna 4.5–5 mm lg., 2.5–3 mm lt.

Terrestrial or lithophytic herb with creeping rhizome and ascending leafy floriferous stem. Rhizome, to 25 cm long and 5 mm wide, light green, thick, at nodes with more or less short rudimentary roots. Floriferous stem ascending, 10-15 cm tall, with (2)3(4) leaves at the base; in upper part covered with 3-4 sterile bracts. Leaves petiolate; petiole and sheath to 5 cm long, 4-6 mm wide at the base; leaf blade thick, slightly succulent, 4-8 cm long, 1.5-3 cm wide, uniform glossy green, with indistinct dark green reticulate network of nerves. Scape to 5 cm long, densely tomentose with soft simple hairs bearing 1-2 (or more?) lax flowers. Floral bracts white, sub-hyaline, with 3 light green nerves, 1.5-2 cm long, 3-5 mm wide, longer than ovary. Ovary 1-1.2 cm long, 2-3 mm broad, densely pubescent with soft simple hairs. Flowers odorless, not widely opening, campanulate, 9-10 mm long.  Sepals white with green tint along mid-vein. Dorsal sepal 8-9 mm long, 5 mm wide. Lateral sepals 8-9 mm long, 3.5-4 mm wide. Petals white, 9-10 mm long, 4-4.5 mm wide, undulate along lateral margin, free at the base, to the apex densely connivent and forming broad hood with the dorsal sepal. Lip white, with yellow spot at the center (yellow distal margin of connivent lobes of mesochile), 10-13 mm long from base to apex of epichile. Hypochile 4-4.5 mm long, 3-4 mm wide; adnate to the column base in basal half; in free distal part with slightly upturned margins; inside on each lateral wall with sea-anemone-like bunch of 6-7 large capitate, arc-divergent glands 1-1.2 mm long; bottom of hypochile with low longitudinal thickening. Mesochile 1.5-2 mm long. Epichile lobes rectangular or sub-quadrate, each 4.5-5.5 mm long, 4-5 mm wide, with 4-6 large irregular dents along distal margin. Column 4.5-5 mm long, 2.5-3 mm wide; stelidia 2-2.5 mm long, rostellar arms 1.8-2 mm long. Anther large, sub-quadrate to indistinctly obovate, 1.8-2 mm long and wide.

Hayata glandulifera Aver.

Etymology. Species name refers peculiar capitate glands inside hypochile.

Flowering time. February.

Ecology. Broad-leaved closed evergreen humid forests on rocky limestone at elev. 400-500 m a.s.l. Creeping terrestrial or lithophytic herb on shady slopes, usually along small forest stream valleys on wet mossy substrate.

Distribution. Vietnam (Phu Tho Province, Tan Son District). Endemic. Fig. 3. New species was found as a very rare plant on the territory of Xuan Son national park (Fig. 3) in small remnant fragment of primary lowland limestone forest. Like recently discovered Vietorchis aurea and Zeuxinella vietnamica (Averyanov and Averyanova, 2003; Averyanov, 2008) discovered species represents element of the lowland highly endangered flora typical for wide limestone areas of northern Vietnam in prehistoric ages and now nearly extinct.